Unit information: Biomechanics and Functional Morphology in 2018/19

Unit name	Biomechanics and Functional Morphology
Unit code	EASCM0024
Credit points	10
Level of study	M/7
Teaching block(s)	Teaching Block 1B (weeks 7 - 12)
Unit director	Professor. Rayfield
Open unit status	Not open
Pre-requisites	n/a
Co-requisites	n/a
School/department	School of Earth Sciences
Faculty	Faculty of Science

Description including Unit Aims

Why are fossil animals shaped in a particular way? How does this relate to their behaviour? In this unit we will examine how biomechanical techniques and inference from living animals shape our understanding of form and function in fossils.

Firstly, students will be introduced to biomechanical principles and techniques, and the issues of adaptation and constraint, before considering their application to problems of fossil animal function. Concepts of basic structural mechanics will be introduced along with an overview of the biology and functional morphology of the musculoskeletal system, including shape and scaling aspects.

The practical application of biomechanics to fossil function will then be considered, focusing on a variety of topics such as feeding and cranial evolution (early tetrapods, lizards and snakes, dinosaurs, mammals), locomotion on land (dinosaurs, hominids and arthropods), in water (plesiosaurs) and in air (insects, pterosaurs, birds.

The unit aims to:

• provide an understanding of the interplay between adaptation and constraint in the form and functional morphology of fossils.

• provide an overview of basic mechanical principles relating to organic structures.

• review and then discuss the applicability of biomechanical techniques in the reconstruction of function in fossils.

• present a series of examples of how biomechanics and inferences from living animals has contributed to our understanding of form and function in extinct animals, in particular focusing on feeding, locomotion, physiology and social interaction in fossils.

Intended Learning Outcomes

On successful completion students will be able to:

• Understand the relationship between form and function in fossils and appreciate why animals may or may not be optimally adapted to particular functions.

• Comprehend the processes by which function can be reconstructed in fossil animals, and evaluate the relative effectiveness of such methods.

• Demonstrate competence in the application of basic biomechanical techniques to functional hypotheses in fossils.

• Possess a broad understanding of how biomechanics has contributed to our understanding of functional evolution throughout the animal kingdom.

Teaching Information

Lectures and practicals

Assessment Information

Coursework (30%). This will be an assessed practical.

2 hour written exam (70%)

Assessment will be completed in accordance with the University Regulations and Code of Practice for Taught Programmes, available online at http://www.bristol.ac.uk/esu/assessment/codeonline.html

Reading and References

Relevant literature will be provided with each lecture and practical class. There are no essential reading texts, but the following are recommended.

• McGowan, C. 1999. A practical guide to vertebrate mechanics. 301pp. Cambridge University Press, Cambridge.
• Biewener, A. A. 2003. Animal Locomotion. 281pp. Oxford University Press, Oxford.
• Thomason, J. J. ed. Functional morphology in vertebrate palaeontology. 277pp. Cambridge University Press, Cambridge.
• Vogel, S. (2013) Comparative Biomechanics, 2nd edition, Princeton University Press, Princeton.

This textbook will be useful for further reading on certain aspects of the course:

• Liem, K. F., Bemis, W. E., Walker, W. F., & Grande, L. 2001. Functional anatomy of the vertebrates. 3rd ed. 703pp. Harcourt, Philadelphia.